Daniel Howell, Ph.D.

Professor of BiologyDaniel Howell

Office: Science Hall 345
Phone: 434-582-2257
E-mail: dhowell@liberty.edu


  • B.S. Old Dominion University
  • Ph.D. Virginia Tech

Postdoctoral Research

  • Duke University Medical Center
  • McGill University 

Courses Currently Taught by Dr. Howell

  • BIOL 213 Human Anatomy & Physiology I
  • BIOL 214 Human Anatomy & Physiology I lab
  • BIOL 215 Human Anatomy & Physiology II
  • BIOL 216 Human Anatomy & Physiology II lab
  • BIOL 313 Clinical Human Anatomy


A native of Lynchburg, Dr. Howell was born at Virginia Baptist Hospital in 1969, the same hospital where today many of his students perform their nursing clinicals. Prior to joining the faculty of Liberty University in 2003, Dr. Howell studied the biochemistry of cystic fibrosis at Duke University Medical Center and then McGill University in Montreal. He currently studies the effects of shoes on foot anatomy and the human gait. Nicknamed the "Barefoot Professor" by his students, Dr. Howell is an avid barefoot runner and hiker and in 2010 published a book (The Barefoot Book) in which he advocates going barefoot more often to improve your health. The book has been translated into several languages and Dr. Howell was featured in a Korean documentary on the benefits of barefoot activities. He has been interviewed by MSNBC, the TODAY Show, Washington Post, Popular Science, Energy Times, Answers magazine, Liberty Journal and many more print, radio and television outlets. Dr. Howell has been invited to speak on the subject of foot health from Portland, OR to London, UK. Married to his wife Carla in 1993, he has three children: Madeleine, Camden and Gillian.

Publications (books, book chapters)

The Barefoot Book: 50 Great Reasons to Kick Off Your Shoes (2010) Hunter House Publishers.

Exploring the Body Machine:  A Laboratory Manual for Exploring Human Anatomy & Physiology, 3rd. (2014). ISBN 978-1-60036-100-5

BIOL 213 lecture guide/BIOL 215 lecture guide, Creation Curriculum. ISBN 978-0-976323-2-7

Hanrahan, J.W., T. Zhu and L.D. Howell. Regulation of CFTR by Protein Phosphorylation, In: The Cystic Fibrosis Transmembrane Conductance Regulator. (2003) Eds. Kevin L. Kirk and David C. Dawson. Plenum Publishers. ISBN 030-6-47837-4

Publications (Journal articles)

Howell, L.D. The transverse arch index: a new method of determining foot arch type from simple footprints. in preparation.

Howell, L.D. The impact of shoes on the human foot arch. in preparation.

Chappe, V., D.A. Hinkson, L.D. Howell, A. Evagelidis, J. Liao, X-B Chang, J.R. Riordan and J.W. Hanrahan. Proc.Natl.Acad.Sci101:390-395, 2004 abstract

Howell, L.D., R. Borchardt, A.M. Kaz, C. Randak and J.A. Cohn. Biochem.J. 378:151-159, 2004 abstract

Derand, R.L., L. Bulteau-Pignoux, Y. Mettey, O. Zegarra-Moran, L.D. Howell, C. Randak, L.J.V. Galietta, J.A. Cohn, C. Norez, L. Romio, J-M. Vierfond, M. Joffre and F. Becq.Am.J.Physiol (Cell Physiol.) 281:C1657-C1666, 2001 abstract

Howell, L.D., R. Borchardt and J.A. Cohn. Biochem.Biophys.Res.Comm. 271:518-525, 2000

McCartney, B., L.D. Howell, M. Potts and P.J. Kennelly. J.Bacteriol 179:2314-2318, 1997

Howell, L.D., C. Griffiths, L. Slade, M. Potts and P.J. Kennelly. Biochemistry 35:7566-7572, 1996

Gordon, A.S., L.D. Howell and V. Harwood. Can.J.Microbiol. 40:408-411, 1994